Abstract
Conservation agriculture involves reduced tillage, permanent soil cover and crop rotations to enhance soil fertility and to supply food from a dwindling land resource. Recently, conservation agriculture has been promoted in Southern Africa, mainly for maize-based farming systems. However, maize yields under rain-fed conditions are often variable. There is therefore a need to identify factors that influence crop yield under conservation agriculture and rain-fed conditions. Here, we studied maize grain yield data from experiments lasting 5 years and more under rain-fed conditions. We assessed the effect of long-term tillage and residue retention on maize grain yield under contrasting soil textures, nitrogen input and climate. Yield variability was measured by stability analysis. Our results show an increase in maize yield over time with conservation agriculture practices that include rotation and high input use in low rainfall areas. But we observed no difference in system stability under those conditions. We observed a strong relationship between maize grain yield and annual rainfall. Our meta-analysis gave the following findings: (1) 92% of the data show that mulch cover in high rainfall areas leads to lower yields due to waterlogging; (2) 85% of data show that soil texture is important in the temporal development of conservation agriculture effects, improved yields are likely on well-drained soils; (3) 73% of the data show that conservation agriculture practices require high inputs especially N for improved yield; (4) 63% of data show that increased yields are obtained with rotation but calculations often do not include the variations in rainfall within and between seasons; (5) 56% of the data show that reduced tillage with no mulch cover leads to lower yields in semi-arid areas; and (6) when adequate fertiliser is available, rainfall is the most important determinant of yield in southern Africa. It is clear from our results that conservation agriculture needs to be targeted and adapted to specific biophysical conditions for improved impact.
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Contents
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Abstract. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
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1. Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5
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2. Materials and methods . . . . . . . . . . . . . . . . . . . . .10
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2.1 Meta-analysis . . . . . . . . . . . . . . . . . . . . . . . . . .10
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2.2 Treatments for the meta-analysis . . . . . . . . . . . . .14
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2.3 Meta-analysis calculations . . . . . . . . . . . . . . . . . .16
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2.4 Rainfall variability and maize yields . . . . . . . . . .17
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2.5 Yield stability analysis . . . . . . . . . . . . . . . . . . . . .18
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3. Results and discussion. . . . . . . . . . . . . . . . . . . . . .18
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3.1 Summary statistics of weighted mean differences. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 18
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3.2 Reduced tillage, continuous maize. . . . . . . . . . . .19
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3.3 No-tillage, maize-legume rotation. . . . . . . . . . . .20
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3.4 No-tillage with mulch, continuous maize. . . . . . .22
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3.5 Effect of mean annual rainfall and rainfall variability. . . . . . . . . . . . . . . . . . . . . . . . . . . .24
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3.5.1 Effect of mean annual rainfall. . . . . . . . . . 24
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3.5.2 Effect of rainfall variability. . . . . . . . . . . . .26
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3.6 Effect of soil texture. . . . . . . . . . . . . . . . . . . . . .28
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3.7 Effect of nitrogen fertiliser input. . . . . . . . . . . . . 29
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3.8 Yield stability analysis. . . . . . . . . . . . . . . . . . . .31
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3.9 Lessons for southern Africa. . . . . . . . . . . . . . . . .33
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3.10 Challenges with long-term experiments. . . . . . .34
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4. Conclusions. . . . . . . . . . . . . . . . . . . . . . . . . . . . .35
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References. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37
1 Introduction
Knowledge of specific crop responses to tillage and surface crop residues as affected by soils, climate and N fertilisation is necessary in the selection of appropriate tillage and crop residue management strategies for improved crop production (Aina et al. 1991). Smallholder agriculture in Southern Africa is characterised by mouldboard ploughing and hand-hoeing that is often thought to lead to land degradation and excessive nutrient losses (Fowler and Rockstrom 2001; Knowler and Bradshaw 2007). To combat this scourge, conservation agriculture is being promoted through reduced tillage, permanent soil cover and crop rotations (FAO 2008). The effectiveness of conservation agriculture for controlling excessive water run-off and soil erosion is well documented (Adams 1966; Alberts and Neibling 1994; Choudhary et al. 1997; Lal 1998; Barton et al. 2004; Scopel et al. 2004), and it is expected that this contribution can be measured in terms of crop yield. Other benefits associated with conservation agriculture include reduction in the input costs for crop production and profit maximisation (Dumanski et al. 2006; Knowler and Bradshaw 2007).
Conservation agriculture emerged in the 1970s mostly in the USA and became an acceptable practice in the USA, Brazil, Argentina, Canada and Australia mainly because of its ability to combat increased soil erosion and land degradation and because of lower fuel costs (Dumanski et al. 2006; Harrington 2008). Conservation agriculture is mostly adopted by large-scale mechanized farmers with the concomitant widespread use of glyphosate for weed control (Derpsch 1999, 2005). Conservation agriculture was developed and adopted widely by farmers in South America mainly because it significantly reduced soil erosion, decreased labour costs and generally led to higher income and a better standard of living for the farmers (Ribeiro et al. 2007; Lahmar 2010).
Implementing conservation agriculture in Africa, particularly the semi-arid regions, presents challenges different from where conservation agriculture originated. In semi-arid regions (300–500 mm annual rainfall), particularly Southern Africa, success of conservation agriculture depends on the ability of farmers to retain crop residues and to ensure adequate weed control (Giller et al. 2009). Farming systems are predominantly mixed crop–livestock systems with low crop productivity and most crop residues are grazed in situ by livestock or transported to the kraal to improve quantity and quality of manure (Murwira et al. 1995; Mapfumo and Giller 2001; Erenstein 2002; Zingore et al. 2007). Rainfall is unimodal and erratic with high variability both within and between seasons, and droughts are common (Challinor et al. 2007). Combined mechanical and hand weeding are the preferred and cheaper weed control methods, and use of herbicides is uncommon (Siziba 2007). Crop rotations are often non-systematic with maize grown continuously for 3–5 years and are aimed at exploiting residual fertility rather than at benefiting the following crops in the rational sequence (Mapfumo and Giller 2001). Fertiliser use is inadequate mainly due to high transaction costs and inefficiencies throughout the production and consumption chain (Quinones et al. 1997). On the other hand, the little fertiliser available is often not the correct type required for various crops and most farmers are not familiar with its correct usage (Sanginga and Woomer 2009).
Manipulating tillage and mulch management to improve water infiltration and reduce water loss from the soil surface in crop fields has potential to substantially improve crop yields and soil conditions in the semi-arid tropics (Hussain et al. 1999; Findeling et al. 2003; Tarkalson et al. 2006; Adekalu et al. 2007). Conventional tillage practices alter soil structure and increase porosity of the upper layer. This increases the initial water infiltration into the soil, but total infiltration is often decreased by subsoil compaction (Aina et al. 1991; Azooz and Arshad 1996; Gómez et al. 1999). Cultivated soils may lose a lot of rainfall as run-off and large amounts of soil through erosion (Duley 1940). Intensive rainfall on bare soil leads to surface sealing and soil compaction, resulting in localised waterlogging and poor soil infiltration (Castro et al. 2006). The mulch component of conservation agriculture controls soil erosion by reducing raindrop impact on the soil surface, decreasing the water runoff rate and increasing infiltration of rainwater (Lal 1989; Barton et al. 2004). Under semi-arid conditions, mulches also play an important role in conservation of soil water through reduced soil evaporation (Scopel et al. 2004). In theory, reduced tillage and surface cover increase soil water available for crop growth by increasing infiltration and by limiting run-off and evaporation losses (Fig. 1). However, mulching is not positive in all circumstances; under continuous rainfall, mulches have little effect on soil water status (Unger et al. 1991). Prolonged dry periods may also cause the benefits of mulching to diminish due to continued evaporation (Jalota and Prihar 1990). Intensive rainfall in mulched fields can cause waterlogging because of reduced evaporation (Araya and Stroosnijder 2010) leading to reduced soil aeration (Cannell et al. 1985).
Interactions between the components of conservation agriculture and their effects on crop yields are complex and often site-specific and long-term experiments are necessary to provide a better understanding. They provide unique information on the sustainability of crop production systems and the interactions between management practices and the broader environment (Powlson et al. 2006). Sustainability is defined as the ability of a system to maintain productivity despite major disturbances such as intensive stress or a large perturbation (Conway 1985; Hansen 1996). Practically, long-term experiments enable observations on changes in crop growth patterns and management effects on slow-moving factors such soil organic matter, which cannot be done in any other way (Jenkinson 1991; Mitchell et al. 1991). They are important for designing cropping systems with high and stable crop yields and low production risk (Raun et al. 1993; Stanger et al. 2008). We analysed maize grain yield data from rain-fed long-term studies on tillage and residue management from semi-arid to sub-humid environments. Maize grain yield is important because it is the staple food crop for most of Southern Africa where it constitutes more than 50% of the diet for most people and can be grown under widely varying rainfall and edaphic conditions (Eicher 1995; Smale 1995; Sileshi et al. 2008). We mainly focused on one of the pillars of sustainable land management, which is to maintain or enhance productivity (Dumanski and Smyth 1994). Crop yield is important because it is the most common and useful parameter used to evaluate the acceptability by farmers of any production practice (Gameda et al. 1997; Abeyasekera et al. 2002).
The objective of this paper was to use data from long-term studies to provide an understanding of the effects of long-term tillage and/or residue retention practices on maize grain yield under contrasting soil textures, crop rotation, N fertiliser input and climate through meta-analysis. An analysis of the relationship between annual rainfall variability and maize grain yield was also carried out using data from southern Africa. This meta-analysis was used to draw major lessons for southern Africa because in this region, there is a strong need for effective soil and water conservation practices to avert the effects of recurrent droughts. Analysing data from other regions provide an indication of the likely impact (ex ante) on food security of promoting reduced tillage and mulch-based cropping practices. It was also intended to understand the interactions between maize yield and rainfall, given its high variability under the climatic conditions of Southern Africa.
2 Materials and methods
2.1 Meta-analysis
Maize grain yield data were obtained from long-term studies (>5 years) on tillage and crop residue management under rain-fed conditions established in semi-arid and sub-humid environments from across the whole world. Treatments had to be from randomised plots with at least three replications. Studies (see Table 1) were obtained from refereed journals, book chapters or peer-reviewed conference proceedings through online searches. Our search was comprehensive including the following keywords and their combinations: conservation agriculture, long-term, reduced tillage, no-tillage, maize yield, corn yield, sub-humid, semi-arid, rain-fed, southern Africa. We also contacted key experts who are working on conservation agriculture. We collected information on climate (mainly rainfall), altitude, soil texture of the experimental site, agronomic management (rate of nitrogen fertiliser applied) as reported by the primary authors (Table 1). These factors were considered to have significant influence on the effect sizes. Data required for the meta-analysis were in the form of treatment mean (\( \overline X \)), its standard deviation (\( {\text{S}}{{\text{D}}_{{\overline X }}} \)) and the number of replicates (n) mentioned in the experimental design. Several authors presented statistical data in different formats such as standard error (\( {\text{S}}{{\text{E}}_{{\overline X }}} \)) and coefficient of variation (\( {\text{CV}}\% \)). These forms were converted to standard deviation (\( {\text{S}}{{\text{D}}_{{\overline X }}} \)) using the following equations: \( {\text{S}}{{\text{D}}_{{\overline X }}} = {\text{S}}{{\text{E}}_{{\overline X }}}\, \times \sqrt {n} \) and \( {\text{S}}{{\text{D}}_{{\overline X }}} = (\frac{{{\text{CV}}\% }}{{100}}) \times \,\overline X \).
Meta-analysis allows quantitative analyses of experimental results reported by other authors and the estimation of effect sizes (Glass 1976; Ried 2000; Rosenburg et al. 2000; Borenstein et al. 2009). The analysis increases the statistical power available to test hypotheses and differences in response between treatments under different environments (Gates 2002; Borenstein et al. 2009). The effect size found in each individual study can be considered an independent estimate of the underlying true effect size, subject to random variation. All studies contribute to the overall estimate of the treatment effect whether the result of each study is statistically significant or not. Data from studies with more precise measurements are given more weight, so they have a greater influence on the overall estimate (Gates 2002). However, meta-analysis has potential weaknesses due to publication bias and other biases that may be introduced in the process of locating, selecting and combining studies (Egger et al. 1997; Noble 2006). Publication bias is the tendency on the part of investigators, reviewers and editors to submit or accept manuscripts for publication based on the direction or strength of the study findings (Dickersin 1990). To overcome these challenges, our searches were carried out online in order to get results from all parts of the world as long as they originated from semi-arid and sub-humid environments. We identified the factors in our analysis such as mean annual precipitation, soil texture and N fertiliser input which could affect the effect sizes and employed the random effects model (Ried 2000).
2.2 Treatments for the meta-analysis
In our analysis, we were interested in treatments that could allow effects of tillage and mulch on maize grain yield to be disaggregated (Table 2). The effect of tillage was analysed by comparing conventional tillage and no-tillage treatments; therefore, conventional tillage was used as the control treatment. No-tillage without rotation was compared with no-tillage with rotation to determine the effect of rotation; thus, no-tillage without rotation was used as the control treatment. Similarly, effect of mulching was analysed by comparing no-tillage without mulch and no-tillage with mulch; therefore, no-tillage without mulch was the control treatment. Moderators of maize yield response were crop rotation, soil texture, mean annual precipitation and N input.
2.3 Meta-analysis calculations
In our analysis, we used the mean difference (Eq. 1.1) in yield between the treatment and control because of its ease of interpretation (Ried 2000). The yield difference is also more relevant when comparing potential gains to required investment and input costs (Sileshi et al. 2008). To obtain overall treatment effects across studies, the differences between treatment and control were weighted (Eq. 1.3). The weight given to each study was calculated as the inverse of the variance (Eq. 1.2). The random effects model was the most appropriate model to calculate effect sizes as it assumed that studies were drawn from different populations, and this could influence the treatment effect. Soil texture, nitrogen input, crop rotation and amount of seasonal rainfall were chosen as covariates and their effect tested on the magnitude of response (mean differences) each with a time component. Due to asymmetry in data distribution between treatments and covariates, conservation agriculture practices (NT, NTR and NTM) were combined together when analysing the effects of covariates. Rainfall was categorised using long-term mean annual of sites to form mean annual precipitation classes as low (<600 mm), medium (600–1,000 mm) and high (>1,000 mm) based on FAO guidelines (Fischer et al. 2001). Soil texture was categorised as clay, sandy, loamy and silt clay loam (Brown 2003), and nitrogen fertiliser input was categorised as low (<100 kg ha−1) and high (>100 kg ha−1) (Osmond and Riha 1996).
2.4 Rainfall variability and maize yields
In sub-Saharan Africa, when sufficient nutrients are available, rainfall variability (both within and across seasons) is the most critical determinant of crop yield (Waddington 1993; Phillips et al. 1998). In this region, 89% of cereal production is rain-fed (Cooper 2004). We evaluated the relationship between maize yield and annual rainfall variability in southern Africa using non-linear regression (Bergamaschi et al. 2007). We used data from three sites with sub-humid climate where long-term conservation agriculture experiments were established in 1988: (1) the Institute of Agricultural Engineering (17°42′ S, 31°06′ E, 1,600 m above sea level and 18 km north of Harare), (2) Domboshawa Training Centre (17°35′ S, 31°10′ E, 1,600 m above sea level and 33 km north of Harare) and (3) Makoholi Research Station (19°34′ S, 30°47′ E, 1,200 m above sea level and 270 km south of Harare). The first site is characterised by deep, well-drained, red clay soils while Domboshawa Training Centre and Makoholi Research Station are characterised by inherently infertile granite-derived sandy soils (Nyamapfene 1991). Both Institute of Agricultural Engineering and Domboshawa Training Centre receive rainfall of about 750 to 1,000 mm/year but Makoholi Research Station receives between 450 and 650 mm/year (Vincent and Thomas 1960; Moyo 2003).
2.5 Yield stability analysis
A stable system shows a small change in response to changes in the environment (Lightfoot et al. 1987). We regarded each tillage practice as a system, and the stability of the system in this study is measured by linear regression of treatment yield against the environmental mean yield; the environmental mean is the average of all the treatments in a given year (Piepho 1998; Hao et al. 2007; Grover et al. 2009). A regression coefficient smaller than one indicates a higher stability (Bilbro and Ray 1976). The regression model is shown in Eq. 1.6:
where y ij is the treatment mean of the ith treatment at the jth environment, μ i is the ith treatment mean in all environments, β i is a regression coefficient corresponding to the ith treatment, u j is an effect of the jth environment and d ij is a random deviation from the regression line (Eberhart and Russell 1966; Piepho 1998).
3 Results and discussion
3.1 Summary statistics of weighted mean differences
Summary statistics showed large variations in maize yield among the treatments across the regions considered (Fig. 2). Reduced tillage with rotation had a positive overall effect on maize yield while reduced tillage (with or without mulch) and continuous maize had negative overall effect on yield compared with the control. Lal (1997) observed that tillage treatments were only significant in three out of eight seasons but maize yield depended more on the amount of rainfall received and its distribution during the season. This observation clearly shows that besides tillage and mulch management, more factors are important for maize yield increases; thus, we explore these factors in the sections that follow.
3.2 Reduced tillage, continuous maize
There was no change in weighted mean differences in maize grain yield over time; therefore, no-tillage had no positive effect on maize yield compared with conventional tillage (Fig. 3). Results showed that in the first 10 years, crops yielded less than the conventional tillage practice. At the beginning of the experiment, reduced tillage practices often resulted in smaller yields than the control, but this was not true for all years. These results are similar to results of Kapusta et al. (1996) who reported no difference in yield between no-tillage and conventional ploughing on poorly drained soils after 20 years of continuous no-tillage. Dam et al. (2005) reported that after 11 years, maize yields were not affected by tillage and residue practices but climate-related differences seemed to have a greater influence on the variation in yields. When residues were completely removed, yield reductions for maize were attributed to decreased soil water storage and excessive surface soil temperatures, especially in climates where conditions of moisture stress occurred during the growing season (Doran et al. 1984). Evidence from Switzerland showed that ploughing could be dispensed under cool moist conditions without loss in yield for crops such as wheat and rape but with maize, no-tillage yielded 10% less than tillage treatments (Anken et al. 2004).
3.3 No-tillage, maize-legume rotation
There was an increase in yield in no-tillage with rotation over no-tillage without rotation as shown by the positive overall weighted mean difference (Fig. 4) in maize–legume rotations. Most of the studies reporting crop yields with rotation showed positive effects in no-tillage systems agreeing with the results of Karlen et al. (1991, 1994a, b), who reported that rotations are likely to produce greater yields across soil fertility regimes. Higher yield for no-tillage in rotation than in monocropping is attributed to a combined effect of multiple factors that include reduced pest infestations, improved water use efficiency, good soil quality as shown by increased organic carbon, greater soil aggregation, increased nutrient availability and greater soil biological activity (Van Doren et al. 1976; Griffith et al. 1988; Hernanz et al. 2002; Wilhelm and Wortmann 2004; Agyare et al. 2006; Kureh et al. 2006). Other authors report that there is often a larger increase in yield in low-yielding environments than in high-yielding environments (Lauer and Oplinger 1996; Porter et al. 1997). The larger yield increase of rotated crops is low-yielding environments means that this production strategy shows promise for most environments in southern Africa. The results of the meta-analysis suggest that rotation should be an integral component of tillage practices for supplying nutrients to maize (Francis and King 1988; Chikowo et al. 2004) and also for breaking pests and disease life cycles as found in other studies (Jordan and Hutcheon 2003; Sandretto and Payne 2007).
3.4 No-tillage with mulch, continuous maize
There was no effect of no-tillage + mulch on yield over the conventional tillage, and after 10 years, there even seems to be strong negative effect (Fig. 5). These results are in contrast with the general belief that conservation agriculture effects emerge in the long term. Results from the Laikipia conservation agriculture project in Kenya show that maize yields were virtually the same under plots managed under conventional tillage and those managed under conservation agriculture (Kaumbutho and Kienzle 2007). Mulch cover associated with no-tillage practices promotes soil water retention (Blevins et al. 1971) and reduces soil temperature (Burrows and Larson 1962) which delays maize emergence and early-season growth. Some authors (Van Doren et al. 1976; Mupangwa et al. 2007) have also found that neither mulching nor tillage practice had a significant effect on maize grain yield on different soil textures and Lal (1997) reported a positive effect of no-tillage + mulch in only three of eight seasons. It has been observed that the effectiveness of mulch is limited in environments with little rainfall (Tolk et al. 1999). The lack of clear benefits on maize grain yield with mulch suggests that it may be better to allocate crop residues as livestock feed instead of keeping it for mulch. Probert (2007) did a modelling exercise using long-term experimental data and concluded that retaining increasing proportions of residues reduces evaporation and run-off but the long-term average yields show only small effects of residue retention on crop yields and the transpiration component of the water balance. Probert (2007) further observed that with no change in transpiration, the reductions in run-off and evaporation must be balanced by increases in drainage. These findings are further supported by a similar modelling exercise using data from the Brazilian Cerrados (Scopel et al. 2004). Vogel (1993) suggested that no-tillage in combination with tied ridging is the most suitable tillage technique for the sub-humid regions because it prevents water-logging and increased root depth, whereas mulching is likely to be the best conservation tillage technique for the semi-arid regions due mainly to reduced topsoil water losses.
3.5 Effect of mean annual rainfall and rainfall variability
3.5.1 Effect of mean annual rainfall
Maize yield was higher with conservation agriculture practices (NT, NTR and NTM) when mean annual precipitation was below 600 mm and lower when mean annual precipitation was above 1,000 mm (Fig. 6). This might be attributed to moisture conservation in low rainfall areas under conservation agriculture and compromised drainage in high rainfall areas. These results agree with Hussain et al. (1999) who reported that yields under conservation agriculture practices were 5–20% lower than under conventional tillage practices in wet years but were 10–100% higher in relatively dry year. Higher crop yield with the conservation agriculture practice than with conventional tillage in a dry year was also reported by Lueschen et al. (1991). Temporal variability in yield is mainly affected by environmental factors with precipitation having the strongest effect (Hu and Buyanovsky 2003; Mallory and Porter 2007; Grover et al. 2009).
3.5.2 Effect of rainfall variability
Variation in total seasonal rainfall across seasons was responsible for major yield fluctuations across treatments in the three experiments of the dataset that were conducted in Zimbabwe (Fig. 7). Rainfall was highly variable across sites and across seasons, at Domboshawa, rainfall varied between 438 and 1,396 mm with a mean value of 823 mm. It caused low yields across all treatments especially in 1989/90, 1991/92 (drought year) and 1996/97. At the Institute of Agricultural Engineering, rainfall ranged between 481 and 1,163 mm with a mean of 889 mm. At Makoholi, rainfall was low but variation between seasons was very high (between 164 and 998 mm) with a mean of 559 mm. In two seasons of contrasting total rainfall, the conventional tillage practice had considerably higher yields than the mulched and reduced tillage treatments, suggesting the absence of benefits of tillage when extreme weather events occur. The low yield during the high rainfall years could be attributed to water-logging that affected nutrient uptake and crop growth (Griffith et al. 1988). The water conservation effect of mulch on maize yield under low rainfall was not observed during the drought of 1991/1992 (Nehanda and Munyati 1999; Moyo 2003). The temporal development of conservation agriculture effects in these three sites seems to be affected more by the amount of seasonal rainfall and soil texture rather than by tillage and mulch management practices. At Domboshawa and Makoholi, both sites characterised by sandy soils, recorded virtually no grain yield during drought years. There are greater chances of conservation agriculture effects developing at the Institute of Agricultural Engineering, which is characterised by a combination of fertile red clay soils and good seasonal rainfall averaging 850 mm in most seasons. The build-up of conservation agriculture effects on sandy soils is a challenge because sandy soils readily lose soil quality during continuous cropping due to compaction, loss of organic matter and acidification (Juo et al. 1996).
3.6 Effect of soil texture
Analysis with soil texture and duration of experiment showed that in clay soils weighted mean differences were mostly negative but were positive in both loam and sandy soils (Fig. 8). There was no significant difference between conservation agriculture treatments (NT, NTR and NTM) and conventional tillage on maize yield on silt clay loams with time. However, there was an improvement in maize grain yield on loamy and sandy soils. Dick and Van Doren (1985) also reported yield reductions of maize associated with no-tillage on heavy clay, very poorly drained soils and suggested crop rotations and use of disease resistant cultivars as possible solutions. However, Van Doren et al. (1976) reported that maize grain yields are insensitive to tillage over a wide range of soil textures, cropping systems, climate conditions and experiment durations as long as equal plant densities and adequate weed control were maintained. The reduction in crop yields on poorly drained soils under conservation agriculture was also reported by Griffith et al. (1988). Increased yields on well-drained soils are attributed to more efficient use of water and improved physical properties (Griffith et al. 1986). Low yields in poorly drained soils are attributed to allelopathy (Yakle and Cruse 1984) and plant pathogens (Tiarks 1977). Kapusta et al. (1996) reported that continuous maize production under no-tillage is most successful on well-drained soil, rather than on either imperfectly or poorly drained soil, especially under wet soil conditions. It has also been suggested that maize monocropping has drastic adverse effects on soil quality and crop yield especially under conditions of low traffic and no-tillage with mulching (Lal 1997). Most soils in southern Africa have biophysical limitations (poor nutrient concentrations, acidity, coarse texture) that limit biomass accumulation; therefore, combinations of legume rotations and mineral nitrogen fertilisation is the most viable option for sustainable agriculture in this region (Chikowo et al. 2004).
3.7 Effect of nitrogen fertiliser input
Nitrogen is often the most limiting nutrient for maize produced in the tropics (Osmond and Riha 1996). At nitrogen fertiliser applications of below 100 kg N ha−1, there were fewer yield advantages of conservation agriculture over conventional tillage, but more yield benefits were obtained with high applications of above 100 kg N/ha (Fig. 9). The results agree with Díaz-Zorita et al. (2002) who reported in a review that maize yields were increased more by nitrogen fertilisation than tillage under sub-humid and semi-arid regions of Argentina. These results show that conservation agriculture practices are input intensive; therefore, improved crop yields under conservation agriculture depend on the ability of farmers to use fertiliser in sufficient quantities and correct proportions. The current average fertiliser use by smallholder farmers in Africa is at 8 kg ha−1 (Groot 2009), and considerable effort is required to improve its use (Sanginga and Woomer 2009). While the fertiliser rates categories considered are quite high and most farmers in Southern Africa cannot afford such rates, fertiliser remains important to alleviate nutrient constraints.
Most crop residues in semi-arid areas are derived from maize, millets and sorghum, which are traditionally known for their poor quality due to high C/N ratios, generally greater than 60 (Cadisch and Giller 1997; Handayanto et al. 1997). Although crop residues are often on the soil surface, there is a greater chance of partial incorporation and decomposition as the season progresses (Parker 1962). The wide C/N ratio and the relatively large amounts of readily decomposable carbon compounds leads to prolonged nitrogen immobilisation by micro-organisms, rendering the nitrogen unavailable for crop growth in the short term (Giller et al. 1997) thus high nitrogen inputs are required when poor quality crop residues are used as mulch.
3.8 Yield stability analysis
There was no treatment effect on stability as a regression between environmental and treatment mean for soil texture (Table 3 and Fig. 10) and for duration of experiment (Table 4 and Fig. 11) with regression coefficients ranging from 0.94 to 1.06 and r 2 values ranging between 0.92 and 0.99. The regression analysis for no-tillage with mulch practice had a smaller regression coefficient in sandy soils showing an advantage of mulch-based systems to optimise moisture availability in soils of poor drainage. Our hypothesis that reduced tillage and residue retention leads to more stable yields was not supported by the data.
3.9 Lessons for southern Africa
Competition for crop residue use, low fertiliser use, non-use of herbicides, labour shortage, erratic rainfall, lack of crop rotations and poor soils combine to offer many challenges for the practice of conservation agriculture among smallholder farmers in Southern Africa (Siziba 2007; Giller et al. 2009). It is clear from the meta-analysis that the success of conservation agriculture in improving crop yields depends on appropriate targeting to climatic and edaphic conditions with adequate inputs (fertiliser and herbicides). Farmers are unlikely to adopt all the conservation agriculture practices and success will not come from the pre-packed technologies alone but from how farmers adapt and apply them depending on resources availability, production objectives (benefits) and biophysical circumstances (Ojiem et al. 2006). In situations of crop–livestock integration where competition for crop residue use is strong, intercropping with grain legumes can be a viable strategy to achieve surface cover because the legume will cover the area between rows of the main crop and help conserve moisture (Scott et al. 1987). In cases were linkages to markets for grain legumes can be secured, legume production can be an excellent opportunity for farmers to increase land size allocated for legumes and improve rotation with main cereal crops. Alternatively planting basins can be an efficient method of moisture conservation if they can be maintained after weeding operations (Mupangwa et al. 2007, 2008).
3.10 Challenges with long-term experiments
Long-term trials are designed to help identify and recommend production systems with beneficial effects on the environment as well as crop productivity across variable environments over time. However, in long-term trials when the cropping system has approached a new equilibrium, it is difficult to attribute effects to particular factors as the interactions between the factors (tillage, mulch, rotation, soil texture and rainfall) involved are so subtle and site-specific that proper experimental designs are required. Sources of variations where crop residues are retained increase as yield varies across seasons to the extent that the effect of mulch will not be explicitly identified. Results from this meta-analysis suggest that yields decline due to continuous monoculture effects and this is more pronounced on sandy soils of low inherent fertility (Lal 1997). These monoculture effects will become more pronounced with time, diminishing the influence of tillage practices on maize yield. Reduction in maize grain yield with continuous maize and no-tillage have been recorded and attributed to unknown underground effects, which need further research (Wolfe and Eckert 1999; Fischer et al. 2002). Well-designed long-term experiments are still desirable across different agro-ecological conditions to unravel the effects of mulch, tillage and rotation on maize grain yield. We propose a simple experimental design (Fig. 12) that we expect can be used to identify the effects of different components. We also propose that the analysis of studies across seasons should take into consideration variability in rainfall to avoid overestimating treatment effects.
4 Conclusions
The factors considered in our analysis covered most of the environments where rain-fed agriculture is practiced and gives us a basis to draw the following conclusions. Positive impacts of moisture conservation on crop yield in soils of poor drainage are likely to occur in low rainfall environments, and maize yielded less in no-tillage without rotation compared with conventional tillage but more when rotation was practised. Results clearly showed that the successful practice of conservation agriculture required high inputs, especially nitrogen fertiliser. Under rain-fed agricultural conditions where total rainfall and its distribution is important for crop production, yield stability analysis results showed that under drought or too much rainfall, no treatments can offset the effects of these extreme conditions. Incentives for abandoning the plough still exist through savings in fuel, labour, and wear and tear of farm implements; however, this needs to be quantified in a separate analysis. Very few studies if any can disaggregate the effects of the three principles (reduced tillage, mulch cover and crop rotation) on maize grain yield; thus, well-designed long-term experiments are still desirable across different agro-ecological conditions to unravel the effects of mulch, tillage and rotation on maize grain yield. Improving maize yields under conservation agriculture in Southern Africa depends on the ability of farmers to practice crop rotation and given that, on average, they plant legumes on 5% of the land, we propose that conservation agriculture be repackaged to reflect the diversity of farming systems and other biophysical and socio-economic considerations for improved impact. Our analyses have shown that success of conservation agriculture in Southern Africa depends on the promotion of other good agronomic practices such as targeted fertiliser application, timely weeding and crop rotations.
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Acknowledgements
We are most grateful and indebted to a number of authors, and we would like to make a special mention to Professor Waren A. Dick, Nithya Janakiraman, Sindhu Jagadamma and Adelaide Moyo-Munodawafa for making available raw data from their long-term studies. Financial support from TSBF-CIAT Harare through the Sub-Saharan Challenge Program’s Conservation Agriculture Taskforce grant is greatly appreciated. Marc Corbeels acknowledges financial support from the European Commission through the CA2Africa Project.
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Rusinamhodzi, L., Corbeels, M., van Wijk, M.T. et al. A meta-analysis of long-term effects of conservation agriculture on maize grain yield under rain-fed conditions. Agron. Sustain. Dev. 31, 657–673 (2011). https://doi.org/10.1007/s13593-011-0040-2
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DOI: https://doi.org/10.1007/s13593-011-0040-2